Ant species dedicated to...

  • Two species dedicated to Prof. Alfred Buschinger.


    Leptothorax buschingeri Kutter, 1967.


    Leptothorax buschingeri Kutter, 1967: 88, figs. 14, 15 (m.). Switzerland. See also: Kutter, 1977: 128.


    Kutter, H., 1967, “Beschreibung neuer Sozialparasiten von Leptothorax acervorum F. (Formicidae).” Mitteilungen der Schweizerischen Entomologischen Gesellschaft, vol. 40: 78-91.

    Kutter, H., 1977, “Hymenoptera, Formicidae.” Insecta Helvetica. Fauna, vol. 6, 1-298.


    Teleutomyrmex buschingeri Lapeva-Gjonova, 2017.

    Tetramorium buschingeri (Lapeva-Gjonova, 2017).


    Teleutomyrmex buschingeri Lapeva-Gjonova, in Kiran, et al. 2017: 151, figs. 3b, 4b, 5b (q.). Bulgaria.

    [Note: Kiran, et al. 2017: 146, retain the “paraphyletic” genus Teleutomyrmex.]


    Kiran, K., Karaman, C., Lapeva-Gjonova, A., Aksoy, V., 2017, “Two new species of the “ultimate” parasitic ant genus Teleutomyrmex Kutter, 1950 from the Western Palaearctic.” Myrmecological News, vol. 25, 145-155.


    Etymology: The species is dedicated to Prof. Alfred Buschinger who has made great contributions to the study of socially parasitic ants.”

  • "Unfortunately Leptothorax buschingeri could never be confirmed as a real species. The type material consists of a series of teratological males." Prof. A. Buschinger.


    But still, the name is accepted and kept in the list of valid species...


    Note on AntWiki: "This species is most probably a junior synonym of Leptothorax acervorum. Only a couple of males from one colony have been described. Having compared the type material with numerous ordinary males of L. acervorum, and with a number of misshaped specimens, I am convinced that L. buschingeri represents nothing but teratological specimens of the presumed host species. A. Buschinger 03:33, 30 November 2012 (EST)."

  • Only 4 ant species are named after Prof. Dr. U. Maschwitz.


    Camponotus (Karavaievia) maschwitzi Dumpert, 2006.


    Camponotus (Karavaievia) maschwitzi Dumpert, in Dumpert, Maschwitz & Weissflog, 2006: 73, figs. 10-13 (w.q.) Borneo.


    Dumpert, K., Maschwitz, U., Weissflog, A., 2006, “Description of five new weaver ant species of Camponotus subgenus Karavaievia EMERY, 1925 (Hymenoptera: Formicidae) from Malaysia and Thailand, with contribution to their biology, especially to colony foundation.” Myrmecologische Nachrichten, vol. 8, 69-82.


    Cladomyrma maschwitzi Agosti, 1991.


    Cladomyrma maschwitzi Agosti, 1991: 306, figs. 5, 8-12, 15, 19, 21, 24 (s.w.q.m.) West Malaysia. See also: Agosti, Moog & Maschwitz, 1999: 11.


    Agosti, D. 1991. Revision of the oriental ant genus Cladomyrma, with an outline of the higher classification of the Formicinae (Hymenoptera: Formicidae). Syst. Entomol. 16: 293-310.

    Agosti, D.; Moog, J.; Maschwitz, U. 1999. Revision of the Oriental plant-ant genus Cladomyrma. Am. Mus. Novit. 3283: 1-24.


    Dolichoderus maschwitzi Dill, 2002.


    Dolichoderus maschwitzi Dill, 2002: 52, figs. III-15, III-23, III-29, III-64 (w.q.m.) Borneo.


    Dill, M., 2002,” Taxonomy of the migrating herdsman species of the genus Dolichoderus Lund, 1831, with remarks on the systematics of other Southeast-Asian Dolichoderus.” Pp. 17-113 in: Dill, M., Williams, D. J., Maschwitz, U., “Herdsmen ants and their mealybug partners.” Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft Frankfurt am Main, vol. 557, 1-373.


    Euprenolepis maschwitzi LaPolla, 2009.


    Euprenolepis maschwitzi LaPolla, 2009: 8, figs. 4A-D (w.) Borneo.


    LaPolla, J. S., 2009, “Taxonomic revision of the Southeast Asian ant genus Euprenolepis.” Zootaxa, vol. 2046, 1-25.


    All the maschwitzi ant species got Maschwitz his name because they wanted to honour him for the behavioural research he did on the ant genera including these species.

  • Who are the people that myrmecologists “use” as name-givers to new ant-species? A few examples from recent tormented ant history…


    Teleutomyrmex schneideri Kutter, 1950.


    Let’s start with the ant Teleutomyrmex schneideri. H. Kutter named that ant after the Swiss Prof. Dr. Otto Schneider-Orelli (1880-1965), his teacher and friend. Prof. Dr. Otto Schneider-Orelli became Doctor in Botany (plants.) in 1905, assistant in botany (grapes for wine) from 1905 till 1913 in Wädenswil and changed to entomology from 1913 till 1917 there. Kept working on insects at the ETH from 1917 on and became professor at the ETH in 1921. In 1928 he became the first director of the Institute of Entomology of the ETH. Kept the functions of professor and director till his retirement in 1950. His most important job was curator of the entomological collection of the ETH in Zurich (1917-1950). He worked as entomologist on Hemiptera as pests in agriculture and forests.

    His father and his son also were botanists (Later, his son became, like his father, an entomologist.).


    Tetramorium schneideri Emery, 1898.


    C. Emery named this ant after Prof. Oskar Schneider, another botanist, from Germany, who had collected the holotype. Couldn’t find more about this Prof. Oskar Schneider.

    When Teleutomyrmex schneideri was placed in Tetramorium it became a junior synonym of Emery’s ant and got another name…


    Teleutomyrmex kutteri Tinaut, 1990.


    Yes, A. Tinaut dedicated this ant to Dr. H. Kutter…


    Tetramorium semilaeve var.kutteri Santschi, 1927.


    In 1927, F. Santschi described the ant and named it after the collector of the animal, H. Kutter. In 2017, the name was synonymised with Tetramorium indocile Santschi, 1927, described in the same paper but four pages earlier…

    When Teleutomyrmex kutteri was placed in Tetramorium it became a junior synonym of Santschi’s ant. Ward et al. didn’t notice that fact and forgot to give it another name…


    So, both ants were named kutteri to honour the same man.

  • Discothyrea stumperi Baroni Urbani, 1977.


    Discothyrea stumperi Baroni Urbani, 1977, 97, figs. 1, 2 (w.) Bhutan.


    Baroni Urbani, C., 1977, “Discothyrea Stumperi n. sp. du Bhoutan, premier représentant du genre dans le Subcontinent Indien (Hymenoptera: Formicidae).” Arch. Inst. Grand-Ducal Luxemb. (n.s.) 37: 97-101.

    De Andrade, M. L., 1998, “First fossil records of the ant genus Discothyrea in Dominican and Mexican amber (Hymenoptera, Formicidae).” Fragm. Entomol. 30: 201-214,


    Temnothorax stumperi (Kutter, 1950).


    Epimyrma stumperi Kutter, 1950: 340, figs. 4-7 (m.) Switzerland. Combination in Myrmoxenus: Schulz & Sanetra, 2002: 162. Kutter, 1951: 153 (w.q.); in Temnothorax: Ward et al., 2014: 15. See also: Stumper & Kutter, 1951: 983; Kutter, 1973: 281.


    Buschinger, A., Ehrhardt, W., Fischer, K., 1981, “Doronomyrmex pacis, Epimyrma stumperi und E. goesswaldi (Hym., Formicidae) neu für Frankreich.” Insectes Sociaux 28: 67-70.

    Kutter, H., 1950, “Über zwei neue Ameisen.” Mitt. Schweiz. Entomol. Ges. 23: 337-346.

    Kutter, H., 1951, “Epimyrma Stumperi Kutter (Hym. Formicid.). 2. Mitteilung.” Mitt. Schweiz. Entomol. Ges. 24: 153-174.

    Kutter, H., 1973, “Beitrag zur Lösung taxonomischer Probleme in der Gattung Epimyrma (Hymenoptera, Formicidae).” Mitt. Schweiz. Entomol. Ges. 46: 281-289.

    Schulz, A., Sanetra, M., 2002, “Notes on the socially parasitic ants of Turkey and the synonymy of Epimyrma (Hymenoptera, Formicidae).” Entomofauna 23(14): 157-172.

    Stumper, R., Kutter, H., 1951, “Sur l'éthologie du nouveau myrmécobionte Epimyrma Stumperi (nov. spec. Kutter).” C. R. Séances Acad. Sci. 233: 983-985.

  • Stumper got two names of ants behind him, Seifert only one:


    Teleutomyrmex seiferti Kiran & Karaman, in Kiran, et al. 2017.

    Tetramorium seiferti (Kiran & Karaman, in Kiran, et al. 2017).


    Teleutomyrmex seiferti Kiran & Karaman, in Kiran, et al. 2017: 148, figs. 3a, 4a, 5a, 6-8 (q.m.) Turkey.

    [Note: Kiran, et al. 2017: 146, retain the paraphyletic genus Teleutomyrmex.].


    Kiran, K., Karaman, C., Lapeva-Gjonova, A., Aksoy, V., 2017, "Two new species of the “ultimate” parasitic ant genus Teleutomyrmex Kutter, 1950 from the Western Palaearctic." Myrmecological News 25: 145-155.

  • Gösswald was a little bit easier, only 3 species names from which only 1 survived!


    Formica goesswaldi Kutter, 1967.


    This taxon is not in use as it is currently considered to be a junior synonym of Formica foreli.


    Formica (Coptoformica) goesswaldi Kutter, 1967: 234, figs. 15-22 (w.q.m.) Germany. Junior synonym of foreli: Seifert, 2000: 544. See also: Kutter, 1977: 284.


    Kutter, H., 1967 [1966], “Einige Ergebnisse weiterer Coptoformica-Studien.” Insectes Soc. 13: 227-240.

    Kutter, H., 1977, “Hymenoptera, Formicidae.” Insecta Helv. Fauna 6: 1-298.

    Seifert, B., 2000, “A taxonomic revision of the ant subgenus Coptoformica Mueller, 1923 (Hymenoptera: Formicidae).” Zoosystema 22: 517-568.


    Leptothorax goesswaldi Kutter, 1967.


    Leptothorax goesswaldi Kutter, 1967: 82, figs. 3-8, 10 (q.) Germany. Kutter, 1977: 129 (m.). Combination in Doronomyrmex: Buschinger, 1981: 215; in Leptothorax: Heinze, 1998: 195 (combination not stated); Bolton, 2003: 270.


    Bolton, B., 2003, “Synopsis and Classification of Formicidae.” Mem. Am. Entomol. Inst. 71: 370pp.

    Buschinger, A., 1974, “Zur Biologie der sozialparasitischen Ameise Leptothorax goesswaldi Kutter (Hym., Formicidae).” Ins. Soc. 21, 133-144.

    Buschinger, A., 1981, “Biological and systematic relationships of social parasitic Leptothoracini from Europe and North America.” P. 211-222 in: Howse, P. E., Clement, J.-L. (eds.), “Biosystematics of social insects.” Systematics Association Special Volume No. 19. London: Academic Press, 346 pp.

    Buschinger, A., 2009, “Social parasitism among ants: a review. (Hymenoptera: Formicidae).” Myrmecological News 12: 219-235.

    Buschinger, A., Klump, B., 1988, “Novel strategy of host-colony exploitation in a permanently parasitic ant, Doronomyrmex goesswaldi.” Naturwissenschaften 75, 577-578.

    Douwes, Per, Abenius, J., Cederberg, B., Wahlstedt, U., 2012, “Steklar: Myror - getingar | Hymenoptera: Formicidae – Vespidae.”

    Heinze, J., 1998 [1995], “The origin of workerless parasites in Leptothorax (s. str.) (Hymenoptera: Formicidae).” Psyche (Camb.) 102: 195-214.

    Kutter, H., 1967, “Beschreibung neuer Sozialparasiten von Leptothorax acervorum F. (Formicidae).” Mitt. Schweiz. Entomol. Ges. 40: 78-91.

    Kutter, H., 1977, “Hymenoptera, Formicidae.” Insecta Helv. Fauna 6: 1-298.

    Ødegaard, F., Olsen, K. M., Staverløkk, A., Gjershaug, J. O., 2015, “Towards a new era for the knowledge of ants (Hymenoptera,Formicidae) in Norway? Nine species new to the country.” Norwegian Journal of Entomology. 62: 80-99.

    Schultz, R., Buschinger, A., 2006, “First Asian record of the parasitic ant, Leptothorax goesswaldi Kutter, 1967 (Hymenoptera: Formicidae).” Myrmecologische Nachrichten. 9: 33-34.


    Epimyrma goesswaldi Menozzi, in Gösswald, 1930.


    This taxon is not in use as it is currently considered to be a junior synonym of Temnothorax ravouxi.


    Epimyrma goesswaldi Menozzi, in Gösswald, 1930: 464, figs. 1-3 (w.q.m.) Germany. [Also described as new by Menozzi, 1931: 41.] Junior synonym of ravouxi: Buschinger, 1982: 352. See also: Stitz, 1939: 186; Kutter, 1973: 281.


    Buschinger, A., 1982, “Epimyrma goesswaldi Menozzi 1931 = Epimyrma ravouxi (André 1896). Morphologischer und biologischer Nachweis der Synonymie (Hym., Formicidae).” Zool. Anz. 208: 352-358.

    Gösswald, K., 1930, “Die Biologie einer neuen Epimyrma art aus dem mittleren Maingebiet.” Zeitschrift für Wissenschaftliche Zoologie 136: 464-484.

    Kutter, H., 1973, “Beitrag zur Lösung taxonomischer Probleme in der Gattung Epimyrma (Hymenoptera, Formicidae).” Mitt. Schweiz. Entomol. Ges. 46: 281-289.

    Menozzi, C., 1931, “Revisione del genere Epimyrma Em. (Hymen. Formicidae) e descrizione di una specie inedita di questo genere.” Mem. Soc. Entomol. Ital. 10: 36-53.

    Stitz, H., 1939, “Die Tierwelt Deutschlands und der angrenzenden Meersteile nach ihren Merkmalen und nach ihrer Lebensweise. 37. Theil. Hautflüger oder Hymenoptera. I: Ameisen oder Formicidae.” Jena, G. Fischer, 428 pp.

  • pisarskii. Formica (Coptoformica) pisarskii Dlussky, 1964: 1034, figs. (w.q.) MONGOLIA. See also: Kupyanskaya, 1990: 204; Seifert, 2000a: 551.

    Dlussky, G. M. 1964. The ants of the subgenus Coptoformica of the genus Formica (Hymenoptera, Formicidae) of the USSR. Zool. Zh. 4 43: 1026-1040 (page 1034, figs. worker, queen described)

    Kupyanskaya, A. N. 1990a. Ants of the Far Eastern USSR. Vladivostok: Akademiya Nauk SSSR, 258 pp. (page 204, see also)

    Seifert, B 2000a. A taxonomic revision of the ant subgenus Coptoformica Mueller, 1923 (Hymenoptera: Formicidae). Zoosystema 22:517-568. (page 551, see also)


    pisarskii. Myrmica pisarskii Radchenko, 1994h: 208, figs. 1-12 (w.q.m.) RUSSIA, MONGOLIA. See also: Radchenko & Elmes, 2010: 216.

    Radchenko, A. G. 1994j. New Palaearctic species of the genus Myrmica Latr. (Hymenoptera, Formicidae). Memorabilia Zool. 48: 207-217 (page 208, figs. 1-12 worker, queen, male described)

    Radchenko, A.G. & Elmes, G.W. 2010. Myrmica ants of the Old World. Fauna Mundi 3: 1-789.


    pisarskii. Strongylognathus pisarskii Poldi, 1994: 187, fig. 1 (w.) ITALY.

    Poldi, B. 1994. Strongylognathus pisarskii species nova (Hymenoptera, Formicidae). Memorabilia Zool. 48: 187-191 (page 187, fig. 1 worker described)


    pisarskii. Temnothorax pisarskii Radchenko, 2004: 115, figs. 7-15 (w.q.m.) NORTH KOREA.

    Radchenko, A. 2004. A review of the ant genera Leptothorax Mayr and Temnothorax Mayr (Hymenoptera, Formicidae) of the Eastern Palaearctic. Acta Zoologica Academiae Scientiarum Hungaricae. 50(2):109-137. (page 115, figs. 7-15 worker, queen, male described)

    Zhou, Shanyi; Huang, Jianhua; Yu, Daojian and Liu, Zhongjian. 2010. Eight New Species and Three Newly Recorded Species of The Ant Genus Temnothorax Mayr (Hymenoptera: Formicidae) From the Chinese Mainland, With a Key. Sociobiology. 56(1):7-26.


    Tetraponera pisarskii

    This taxon is not in use as it is currently considered to be a junior synonym of Tetraponera modesta.

    pisarskii. Tetraponera pisarskii Radchenko, 1997g: 480, figs. 1, 2 (w.) KOREA. Junior synonym of modesta: Ward, 2001: 616.

    Radchenko, A. G. 1997h. Tetraponera pisarskii sp. nov. - the first native member of subfamily Pseudomyrmicinae from Palaearctic Asia. Ann. Zool. (Warsaw) 47: 479-480.(page 480, figs. 1, 2 worker described)

    Ward, P. S. 2001. Taxonomy, phylogeny and biogeography of the ant genus Tetraponera (Hymenoptera: Formicidae) in the Oriental and Australian regions. Invertebr. Taxon. 15: 589-665 (page 616, Junior synonym of modesta)


    pisarskii. Tetramorium pisarskii Radchenko & Scupola, 2015: 226, fig. 5 (w.) AFGHANISTAN.

    Radchenko, A.G. and Scupola, A. 2015. Taxonomic revision of the strativentre species group of the genus Tetramorium (Hymenoptera, Formicidae). Vestnik zoologii. 49:219–244.


    6 species named after B. Pisarski, 5 still valid, 1 not valid.

  • kutteri, 11 ants named after H. Kutter, 6 still valid, 4 as synonyms and 1 unavailable.


    Camponotus kutteri

    kutteri. Camponotus (Myrmamblys) kutteri Forel, 1915a: 40 (s.w.q.m.) INDONESIA (Sumatra).

    Forel, A. 1915a. Fauna Simalurensis. Hymenoptera Aculeata, Fam. Formicidae. Tijdschr. Entomol. 58: 22-43 (page 40, soldier, worker, queen, male described)


    Temnothorax kutteri

    kutteri. Chalepoxenus kutteri Cagniant, 1973: 148, figs. 1-4 (w.q.m.) FRANCE. Combination in Temnothorax: Ward et al., 2014: 15.

    Buschinger, A. (2009) Social parasitism among ants: a review. (Hymenoptera: Formicidae). Myrmecological News 12: 219-235.

    Cagniant, H. 1973b. Description et représentation des trois castes de Chalepoxenus kutteri (nov. sp.) (Hyménoptères Formicidae - Myrmicidae). Insectes Soc. 20: 145-156 (page 148, figs. 1-4 worker, queen, male described)

    Ward, P.S., Brady, S.G., Fisher, B.L. & Schultz, T.R. 2015. The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology, 40: 61-81.


    Crematogaster kutteri

    kutteri. Crematogaster kutteri Viehmeyer, 1924b: 314 (w.) AUSTRALIA. Combination in C. (Crematogaster): Bolton, 1995b: 166

    Viehmeyer, H. 1924c. Formiciden der australischen Faunenregion. (Fortsetzung). Entomol. Mitt. 13: 310-319 (page 314, worker described)

    Bolton, B. 1995b. A new general catalogue of the ants of the world. Cambridge, Mass.: Harvard University Press, 504 pp.


    Leptothorax kutteri

    kutteri. Leptothorax (Mychothorax) kutteri Buschinger, 1966a: 327, figs. 1, 2, 3 (q.m.) GERMANY. Combination in Doronomyrmex: Buschinger, 1981: 215; in Leptothorax: Heinze, 1998: 195 (combination not stated); Bolton, 2003: 270. See also: Kutter, 1967b: 85; Kutter, 1977c: 130; Buschinger, 1972: 169.

    Blatrix R., Lebas C., Wegnez P., Galkowski C., Buschinger A., 2013. New data on the distribution of Leptothorax pacis and L. kutteri, two very rare parasitic ants, and confirmation of the presence of L. gredleri in France (Hymenoptera, Formicidae). Revue de l’Association Roussillonnaise d’Entomologie, 22, 85-91.

    Bolton, B. 2003. Synopsis and Classification of Formicidae. Mem. Am. Entomol. Inst. 71: 370pp (page 272, Combination revived in Leptothorax)

    Buschinger, A. 1966a [1965]. Leptothorax (Mychothorax) kutteri n. sp., eine sozialparasitische Ameise (Hymenoptera, Formicidae). Insectes Soc. 12: 327-334 PDF (page 327, figs. 1, 2, 3 queen, male described)

    Buschinger, A. 1972a. Kreuzung zweier sozialparasitischer Ameisenarten, Doronymyrmex pacis Kutter und Leptothorax kutteri Buschinger (Hym., Formicidae). Zool. Anz. 189: 169-179 (page 169, see also)

    Buschinger, A. 1981. Biological and systematic relationships of social parasitic Leptothoracini from Europe and North America. Pp. 211-222 in: Howse, P. E., Clement, J.-L. (eds.) Biosystematics of social insects. Systematics Association Special Volume No. 19. London: Academic Press, 346 pp. (page 215, Combination in Doronomyrmex)

    Buschinger, A. (2009) Social parasitism among ants: a review. (Hymenoptera: Formicidae). Myrmecological News 12: 219-235.

    Heinze, J. 1998 [1995]. The origin of workerless parasites in Leptothorax (s. str.) (Hymenoptera: Formicidae). Psyche (Camb.) 102: 195-214 (page 195, Returned to Leptothorax)

    Kutter, H. 1967c. Beschreibung neuer Sozialparasiten von Leptothorax acervorum F. (Formicidae). Mitt. Schweiz. Entomol. Ges. 40: 78-91 (page 85, see also)

    Kutter, H. 1977c. Hymenoptera, Formicidae. Insecta Helv. Fauna 6: 1-298 (page 130, see also)

    Ødegaard, F., K.M. Olsen, A. Staverløkk, and J. O. Gjershaug. 2015. Towards a new era for the knowledge of ants (Hymenoptera, Formicidae) in Norway? Nine species new to the country. Norwegian Journal of Entomology. 62:80-99.


    Myrmecina kutteri

    This taxon is not in use as it is currently considered to be a junior synonym of Myrmecina graminicola.

    kutteri. Myrmecina kutteri Forel, 1914b: 1 (w.) ITALY. [Also described as new by Forel, 1915d: 21 (w.q.m.).] Subspecies of graminicola: Emery, 1916b: 171. Junior synonym of graminicola: Brown, 1951: 106.

    Brown, W. L., Jr. 1951. New synonymy of a few genera and species of ants. Bull. Brooklyn Entomol. Soc. 46: 101-106 (page 106, Junior synonym of graminicola)

    Emery, C. 1916a [1915]. Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bull. Soc. Entomol. Ital. 47: 79-275 (page 171, Variety of graminicola)

    Forel, A. 1914b. Deux nouveautés myrmécologiques. Yvorne, Switzerland: published by the author, 1 p. (page 1, worker described)

    Forel, A. 1915d. Fauna insectorum helvetiae. Hymenoptera. Formicidae. Die Ameisen der Schweiz. Mitt. Schweiz. Entomol. Ges. 12(B Beilage: 1-77 (page 21, [Also described as new by Forel, 1915d: 21 worker, queen, male described)


    Tetramorium kutteri

    kutteri. Teleutomyrmex kutteri Tinaut, 1990b: 202, figs. 1-3, photos. 1-2 (q.m.) SPAIN. Combination in Tetramorium: Ward et al., 2014: 16.

    Buschinger, A. (2009). Social parasitism among ants: a review. (Hymenoptera: Formicidae). Myrmecological News 12: 219-235.

    Kiran, K., Karaman, C., Lapeva-Gjonova, A. & Aksoy, V. 2017. Two new species of the “ultimate” parasitic ant genus Teleutomyrmex Kutter, 1950 from the Western Palaearctic. Myrmecological News 25: 145-155.

    Tinaut, A. 1990b. Teleutomyrmex kutteri, spec. nov. A new species from Sierra Nevada (Granada, Spain). Spixiana 13: 201-208 (page 202, figs. 1-3, photos. 1-2 queen, male described)

    Ward, P.S., Brady, S.G., Fisher, B.L. & Schultz, T.R. 2014. The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology, 40: 61-81.


    Messor semirufus kutteri

    This taxon is not in use as it is currently considered to be a junior synonym of Messor semirufus.

    kutteri. Messor semirufus var. kutteri Santschi, 1934d: 275 (w.) ITALY. Junior synonym of semirufus: Baroni Urbani, 1974: 227.

    Baroni Urbani, C. 1974a. Studi sulla mirmecofauna d'Italia. XII. Le Isole Pontine. Fragm. Entomol. 9: 225-252.

    Santschi, F. 1934d. Fourmis d'une croisière. Bull. Ann. Soc. Entomol. Belg. 74: 273-282.


    Myrmica schencki kutteri

    This taxon is not in use as it is currently considered to be a junior synonym of Myrmica schencki.

    kutteri. Myrmica schencki var. kutteri Finzi, 1926: 111 (w.m.) SWITZERLAND. Junior synonym of schencki: Bernard, 1967: 118.

    Bernard, F. 1967a [1968]. Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp.

    Finzi, B. 1926. Le forme europee del genere Myrmica Latr. Primo contributo. Boll. Soc. Adriat. Sci. Nat. Trieste 29: 71-119


    Tetramorium semilaeve kutteri

    This taxon is not in use as it is currently considered to be a junior synonym of Tetramorium indocile.

    kutteri. Tetramorium semilaeve var. kutteri Santschi, 1927a: 57 (w.) SWITZERLAND.

    Subspecies of semilaeve: Novák & Sadil, 1941: 85 (in key); Bolton, 1995b: 410.

    Junior synonym of indocile: Wagner, et al. 2017: 116.

    Bolton, B. 1995b. A new general catalogue of the ants of the world. Cambridge, Mass.: Harvard University Press, 504 pp.

    Novák, V.; Sadil, J. 1941. Klíc k urcování mravencu strední Evropy se zvlástním zretelem k mravencí zvírene Cech a Moravy. Entomol. Listy 4: 65-115

    Santschi, F. 1927b. A propos du Tetramorium caespitum L. Folia Myrmecol. Termit. 1: 52-58 (page 57, worker described)

    Wagner, H.C., Arthofer, W., Seifert, B., Muster, C., Steiner, F.M. & Schlick-Steiner, B.C. 2017. Light at the end of the tunnel: Integrative taxonomy delimits cryptic species in the Tetramorium caespitum complex. Myrmecological News 25: 95-129.


    Camponotus kutterianus

    kutterianus. Camponotus kutterianus Baroni Urbani, 1972: 132, figs. 1-3 (w.) CUBA.

    Baroni Urbani, C. 1972. Studi sui Camponotus (Hymenoptera, Formicidae). Verhandlungen der Naturforschenden Gesellschaft in Basel 82:122-135.


    Strongylognathus huberi st. cecconii var. kutteri Santschi, 1927a: 58 (q.) ITALY (Sicily). Unavailable name; material referred to destefanii by Baroni Urbani, 1964b: 54.

    Santschi, F. 1927b. A propos du Tetramorium caespitum L. Folia Myrmecol. Termit. 1: 52-58.

    Baroni Urbani, C. 1964b. Studi sulla mirmecofauna d'Italia. II. Formiche di Sicilia. Atti Accad. Gioenia Sci. Nat. (6) 16: 25-66.

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